(Exciting Comics 005, 1940)
Stretching the definition of "super-villain" to the very limits here: the Tankonaut is a superweapon developed by US arms manufacturer M.V. Gage under contract to a man who is very clearly meant to be Hitler but is referred to only as the Dictator. Whether this would have qualified as treason in mid-to-late 1940 is beyond me but the next part, where the Tankonaut is tested on US targets (including Washington DC!) extremely is.
Not only does the Tankonaut have an excellent name (100% the only reason it made the grade as an entry here, even under my already very loose policies about super-vehicles) but it has a neat gimmick: it isn't a tank at all! Rather, it is a big beefy truck that carries around its own armour plating and weaponry for rapid deployment/ quick and sneaky escape! It's a neat variation on the old "drive the crime vehicle into the back of a truck" scheme for the truck to actually be the crime vehicle.
Unfortunately for M.V. Gage and the Tankonaut, John Thesson, the Son of the Gods, is on the case. He discovers the Tankonaut's secret identity of A Truck by the simple expedient of flipping it over with a long pole. As usual, once the super vehicle has been disabled the non-super humans are quickly rounded up.
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